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Houston writer Bryan Washington to receive Gaines award |
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Arizona Wildcats at Washington Huskies 01/30/20 Free Pick amp Prediction |
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Internship And Study Abroad Provider business for sale Washington DC |
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From Italy to Washington DC: Enrica Consultant at OSA |
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The Washington Center |
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Alberta patří Calgary! Nashville uspěl na ledě Washingtonu Hertl se zranil |
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NCWV Airport officials discuss adding second daily round-trip flight to Washington DC |
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Séjour du Chef de l’Etat à Washington : Patrice Talon échange avec le Vice-président de la Banque Mondiale et avec la diaspora béninoise |
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Top 20 Best Plumbers Near Me in Washington |
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En Washington el Gobierno defiende actuación en Santurbán |
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Washington State Attorney General Bob Ferguson Looking Into Decision to Close National Archives in Seattle |
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Pot Politics: The Marijuana Business enterprise Concerns Washington |
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Tennessee Lawmaker Introduces Bill Labeling CNN Washington Post as ‘Fake News’ |
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Alex Ovechkin honors Kobe Bryant by wearing No 24 jersey before Washington Capitals game |
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Washington state launches law enforcement hiring initiative |
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University of Washington |
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Nashville Predators vs Washington Capitals |
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Patrice TALON à Washington : Le point de ses rencontres et activités |
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Washington County |
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5 Things Washington: Health policy update Coronavirus Sen Manka Dhingra |
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Republican State Rep Introduces Resolution to Brand CNN and Washington Post as ‘Fake News’ and ‘Media Wing of Democratic Party’ |
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Toko Daging Washington Jual Daging Halal untuk Dukung Mahasiswa Muslim |
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Which Iconic Sci-Fi Character Is Immortalized As A Grotesque On The Washington National Cathedral |
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Amblin Television amp Alex Gibney To Adapt The Washington Post’s ‘Afghanistan Papers’ Into Limited Documentary Series amp Drama |
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PBJ Marketing A Digital Marketing Agency Located in Washington DC and New York City |
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CJ Elleby hits winner securing Washington State’s dramatic 67-65 win over Arizona State |
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Important Speeches and Documents in the Formation of America: George Washington |
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Washington DUI Process From Traffic Stop to First Appearance |
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Washington Kirkwood |
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Master’s Conference – Washington DC – 14 years |
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Charlotte Hornets vs Washington Wizards |
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Washington Enacts Law Creating New Paint Recycling Program |
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IITA Washington University in St Louis to jointly tackle poverty and hunger |
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WASHINGTON DC |
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Focus is on fun as Washington gets set for Arizona schools |
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Embassy in Washington |
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Washington Library |
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En séjour à Washington: Talon positionne le Bénin au cœur de la Banque Mondiale |
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Patrice Talon a rencontré Mike Pompeo à Washington |
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Washington Post reverses suspension of reporter who tweeted about Kobe Bryant |
2 |
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Bennett: Picks against the spread for Iowa-Maryland Arizona-Washington and other top games |
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Washington DC published |
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WSU Washington well represented in online architecture database |
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Dimambro testifies he didn’t harm 2-year-old Damian in Washington Township home |
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Oaklawn Park Barn Notes: Taraz Ready for Saturay’s Martha Washington Stakes |
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Cowles Co Secures Financing for Washington Community |
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Washington Township’s Woodmont Townsquare reaches full occupancy |
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Accredited Dual Diagnosis Treatment in Washington State |
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16 tested for coronavirus in Washington |
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The Dankest Cannabis Cupcakes in Washington: BONG APPÉTIT… |
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Climate Change Activists Attempt to Shut Down Washington DC |
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The Most “Instagrammable” Photo Spots in Washington DC |
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Local Washington Affiliates |
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19:00 – Washington Wizards Chicago Bulls – NBA Basketball |
2 |
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Alkitab Ibrani Berusia 1000 Tahun di Publikasikan di Museum of the Bible Washington DC |
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Dumpster Rental in Washington DC |
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19:00 – Cleveland Cavaliers Washington Wizards – NBA Basketball |
2 |
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Washington Capitals new service dog Scout makes debut |
2 |
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Washington DC Sports Betting App delayed to March |
2 |
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Newly Obtained Documents Reveal Washington Department of Fish and Wildlife Side-Stepped Non-Lethal Actions Before Wiping Out Wolf Packs |
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Armed robbers raid jeweller in Washington Mall |
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Southwest Washington Fairgrounds |
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Keep it Clean Washington DC – Dumpster Rental Services in DC |
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Washington DC Regional Headquarters |
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Walter E Washington Convention Center |
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Throwback Thursday: The birth of Washington-Kosciusko School in Winona |
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CarBelles SoulBounce and Black Girls Drive -Kick Off Washington DC Auto Show with Kia and Mazda |
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Washington DC Tax amp IRSHot Topics |
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Leadership Washington Business amp Industry Day |
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An Exclusive Interview With Asa Palagi an Independent Candidate for Governor of Washington |
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Washington DC to Dublin Non-Stop for 416 |
2 |
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Amber Athey joins as The Spectator’s Washington editor |
2 |
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Washington College |
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The ULTIMATE Washington DC Travel Guide |
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Washington sees surge of foster kids placed in hotels while they wait for treatment |
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Register today to be part of the Greek American business community Hermes Expo is hosting its 29th Business and Educational Expo in Chicago Washington Philadelphia and New York this coming April |
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Washington Community Schools |
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Theatre Review: ‘Bloomsday’ Washington Stage Guild |
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Washington Evening Journal |
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Homosexual Friend Dating Solution Washington Dc |
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Sheriff: Suspected kidnapper at large after high-speed chase crash in Washington Co |
2 |
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Ford traveling to Washington DC to unveil new trade strategy |
2 |
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Washington County coalition brings concerns to state legislators |
2 |
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Washington Post Reporter Felicia Sonmez amp Others Smear Kobe Bryant Shortly After His Death |
1 |
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At Washington Law Center… we fight you win |
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Advocacy Alert: DJ Jaffe at White House DJ Jaffe on Fox Local Solutions Fact Sheet Money available for your state hospital…John Stossel interviews DJ Jaffe…Our controversial TEDTalk…Our Washington Post oped…How to reduce the mental illness to jail pipeline… Fed Serious Mental Illness Committee Reports on Needs of Seriously Mentally Ill… |
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Here’s how Washington State’s nudge for state park donations works via its web site |
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‘Paint in the Park’ at Booker T Washington National Monument |
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City: Undisclosed State: md County: Washington |
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It’s Not Just Washington State Iranians Are Stopped At Borders All Over The Country |
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Prom Limousine Service in Washington DC Virginia |
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Washington Greene responders get 850K in grants |
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Washington Investiture de Trump « aujourd’hui nous rendons le pouvoir au peuple » |
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BizTimes Media’s Washington County 2035 |
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Timuçin Sahin Trio feat Reggie Washington amp Sean Rickman |
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Michael Washington |
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About Washington Regional |
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300 s Washington avenue 3rd floor Greenville MS 38701 |
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Hillman Hall at Washington University |
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The Arctic Next Door: Mount Washington |
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Mistakes with Your Kitchen Remodel Design You Must Avoid in VIRGINIA MARYLAND and WASHINGTON DC AREA WİTH EIGHT SHOWROOMS |
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The Jordan Schnitzer Museum of Art at Washington State University |
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How To Find Affordable Health Insurance In Washington |
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Essentials of NLP June 12-14 2020Cascade HypnosisBellingham Washington |
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Northern Virginia Montgomery County MD and Washington DC |
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Join us at events throughout Washington |
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Experience the world of wineries wine events wine bars and tasting rooms across Washington wine country Let’s begin |
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Coronavirus may have originated in lab linked to China’s biowarfare program The Washington Times – Sunday January 26 2020gt Người ta nghi ngờ siêu vi khuẩn corona có nguồn gốc từ các phòng nghiên cứu võ khí sinh học của Trung Quốc … |
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Washington DC and Alexandria Ghost Tours |
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Two articles from the Washington University Student Life Newspaper |
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George Washington Carver on Ego and Self |
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Story Time at Washington Park |
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Mr Baron Goes to Washington |
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The CPC – Seattle Washington Dispensary |
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WASHINGTON MONUMENT amp DC HIGHLIGHTS TOUR |
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Ο Trump στην πορεία για τη ζωή στην Washington |
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Washington DC to Paris for Just 432 |
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Mt Washington adventure It lies in wait for the opportunity to come out and play |
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Kerry Washington Discusses Her Evolution as an Actor an Activist and an American |
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Grand Hyatt Washington DC Learn more gtgt |
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UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON DC FORM 10-Q |
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Lovely Bride: Washington DC |
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Western Washington 48 magnitude earthquake! |
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Washington Redskins Football Dog Toy |
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Management and Program Analyst Freedom of Information Act FOIA US Department of Homeland Security Office of the Chief Privacy Officer Washington DC |
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Oficina Antofagasta Jorge Washington 2701 Antofagasta Región de Antofagasta |
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At a time when the messages we give and get have so many more ways to be misconstrued and potentially damaging a book that takes apart our language becomes almost vital to our survival as friends -THE WASHINGTON POST |
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Washington Dc |
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Welcome ToThe Washington Metro Area |
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No te dejes manipular: USA monta show de un ovni en pleno Washington! |
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Diane Johnson Washington DC |
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Pinterest Bans Misinformation About Voting and the Census washingtonpostcom 25 |
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‘Morning Joe’ Anchors Navigate While ‘Washington Is on Fire’ time-clock Created with Sketch 10:00AM PST |
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Kim Jong-Un Son message codé pour Washington |
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Washington Post’s Wonkblog Pundit of the Year |
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Séisme à Washington : les démocrates lancent une procédure de destitution contre Trump |
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New Daily Flights between Washington Dulles and TVC starting June 2020 |
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Shipping Container Home – The SurfShack Hartman Kable – Grayland Washington |
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Zvykl jsem si tady říká o Washingtonu Radko Gudas Je rád že nemusí bránit Ovečkina |
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Washington Goncalves de Tian |
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Washington Capitals Team Plaque 12″ x 40″ |
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De hecho cuando Estados Unidos aceptó a Libertas como su símbolo e identidad su capital Washington DC ya era famosa por sus muchos burdeles Un verdadero gobierno cristiano nunca aceptaría Libertas Un gobierno con su capital lleno de burdeles la recibiría calurosamente |
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Gov Stitt to join Trump administration officials in Medicaid announcement in Washington DC |
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The Washington Connection and Third World Fascism |
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George Washington Thanksgiving Historical Proclamation 1789 |
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A FUTURE HOME: TWO DOZEN STUDENTS EXPLORE 377 WASHINGTON UNDER that Inspire |
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The Star Cities are Start Cities – Join the Event in One of These Cities and Ride East to Washington DC with us |
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Washington Law Center has the experience and resources you need If you are seeking a personal injury lawyer or a Labor and Industries LampI attorney in the state of Washington call us today |
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Washington Wizards Play WIFFLE Ball At Practice |
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Denzel Washington Wife 2018 |
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Welcome to Washington County Maryland! |
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Middle East Investment Initiative2300 N Street NW Washington DC 20037 |
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Licensing Outdoor amp Nature-Based Preschools In Washington State and Beyond |
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Washington DC Agenda Published |
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5 BEST Museums in Washington DC |
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Explore the islands of Washington State’s San Juan archipelago on this kayaking and camping adventure 48 Hours Left To Book |
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Washington DC to Mumbai |
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Servicing Baltimore Washington DC and Northern VA |
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Washington Advanced |
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Judy Martin elected Chair of Washington’s Global Ageing Network GAN |
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Add your Business to Washington Yellow Pages for Free |
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New report highlights needs and experiences of independent contractors in Washington State |
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WELCOME TO WorkSource WASHINGTON |
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Questions and Answers about a Novel Coronavirus in Washington |
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Redefining what’s possible for education in Washington state |
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Advocates Law Group: Washington |
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A Taste of Washington |
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Premier Auto Group is the Automotive Home for Mercedes-Benz Toyota Lexus Ford and Chevrolet Drivers in Turnersville Washington Township Glassboro and Lindenwold |
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Coalition Statement on Reports of Mass Detentions of Iranians and Iranian Americans at the Washington/Canada Border |
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One of George Washington’s swords |
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Vietnam : Un témoignage historique de la sauvagerie de Washington 21st century wire |
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This daily email features the latest news for Washington County Greene County and the Mon Valley |
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Ideal Home Care Agency Serving Washington State |
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2020 Washington Mead amp Cider Cup Homebrew Competition |
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Micro-Mobility Leader Helbiz Launches e-Bikes in Washington DC – Associated Press |
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Washington Independent Review of Books |
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Trump returns to Washington to face a pair |
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Washington Supreme Court: When a juror of color reports experiencing racial bias the court must investigate |
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Washington DC Temple: A Beautiful Sight |
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Robert Laszewski Washington DC |
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CSA Joins AFSA To Lobby For Educators in Washington DC |
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Washington introduces sports betting bills |
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Masters in Computer Science at Washington University in StLouis Missouri USA |
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So you’ve found your favorite Mercedes-Benz Toyota Lexus Ford or Chevrolet: now what Premier Auto Group Proudly Serving Washington Township Glassboro and Lindenwold Makes Car Financing Simple for Drivers in Turnersville |
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Seattle Washington USA Downsizing Online Auction – 46th Avenue So |
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PhotoSecretsWashington at Powell’s |
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Washington Park Elementary seeks donations for kindergarten biking program |
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Washington Post suspendió a una periodista por un tweet contra Kobe Bryant |
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NEW PODCAST: Blake Mazza – Washington State Kicker |
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Memorable Quotations from George Washington Kindle Book |
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Thompson Readies for Opening in Washington DC |
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WASHINGTON DC 2014 |
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Charlotte Hornets Vs Washington Wizards |
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Guided Washington DC Bus Tours by USA Guided Tours |
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Pedestrian Cheryln Ninette Bell 49 killed in hit-and-run collision on Suitland Parkway in Washington DC |
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Newspaper industry loses one of its last big backers in Buffett who delivered Washington Post as a kid |
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Washington Video Production |
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Washington to Hyderabad |
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Lambert – Washington |
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Washington State Road Map |
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Washington Post: As impeachment vote looms Pompeo’s Ukraine trip is a high-risk high-reward venture |
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Trump e von der Leyen vertice a Washington per fermare i dazi americani |
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If you live work worship or attend school in Smyth Wythe or Washington counties you and your family members are eligible to join Mountain Empire FCU Organizations located in these counties also qualify for membership |
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Stop in and visit us today! We are located at 5001 New Hampshire Avenue NW Washington DC 20011 Phone: 202-726-3100 Fax: 202-291-5259 |
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Ona Judge – Slave and Martha Washington’s Personal Servant |
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Wild ending leaves Arizona State with rough loss to Washington State |
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Gantz heads to Washington for peace plan talks with Trump |
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Washington Communities |
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Washington University School of Medicine USA |
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George Washington |
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Washington Post: It’s important to take service members’ brain injuries seriously |
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New North Concourse at Ronald Reagan Washington National Airport DCA |
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Washington se dit « indigné » par une nouvelle attaque contre une base en Irak |
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Kerry Washington Goes For Plaid |
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DAVID PICKERAL Mobility and Smart Cities Expert Washington |
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2018 Conventional Loan Limits In Washington State |
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Tacoma Washington |
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Juan Carlos I el espía de Washington |
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BWI Airport to Washington DC Shuttle IAD Airport Shuttle – Washington Dulles DCA Airport Shuttle – Ronald Reagan National Airport |
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So far under California’s new privacy law firms are disclosing too little data – or far too Washington Post |
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Boris Johnson en búsqueda del equilibrio justo con Washington y Bruselas EXCLUSIVO PARA SUSCRIPTORES |
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Washington DC Video Production |
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Rent Distinctive Courtroom Reporting Washington DC Service |
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Upcoming Vaccine Congress Washington |
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Puppetry festival to open in Washington |
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Washington DC Mayor Muriel Bowser endorses Mike Bloomberg |
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La música debe crecer: Kamasi Washington |
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Europe’s Dangerous Irrelevance in Washington and the Middle East |
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Washington column for Jan 16 |
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Quand le Maroc donnait l’exemple du respect de la légalité lors de la Guerre civile américaine Washington – La revue scientifique américaine ”Smithsonian Magazine” revient dans un récent article sur un épisode méconnu de la Guerre de sécession aux Etats-Unis Baptisé affaire DeLong ou diff |
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HistoryR to Premiere Epic Three-Night Miniseries Event “Washington” Presidents’ Day Weekend Beginning Sunday February 16 at 8PM VIDEO |
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Webinar Recap: Washington Outdoor Preschool Pilot |
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Washington Sign Up |
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Timothy Washington: Citizen/Ship |
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Hagerstown amp Washington County Maryland |
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Manuel Sutherland: Lo más duro para Maduro sería salir en TV y decir “vamos a dolarizar formalmente y que viva Washington” |
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Solution-Focused Washington DC Tax Attorney |
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Western Washington University |
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Over 20 Destinations in the Washington State |
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Proposed Gambling Law Revision Aimed at Protecting Washington State Gaming Industry |
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TH AVE NE BELLEVUE WASHINGTON |
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Brian Blase on C-SPAN’s Washington Journal |
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Trinity HealthShare programs are not available in AK CO CT HI MA MD ME MT ND NH OR PA PR SD TX VT WA WY or Washington DC Limitation subject to change without prior notice Due to regulatory limitations regarding compensation Trinity HealthShare programs will no longer be sold in Massachusetts or Pennsylvania |
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WITH WASHINGTON DC Sample 3-Day from 263 |
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New Pakistan Ambassador Brings Frank Talk to Washington Relations |
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Featuring → Your Washington Spot |
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Fox 5 Washington DC Live Streaming |
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Washington DC’s top speakers bureau for keynote speakers motivational speakers political speakers leadership speakers business speakers moderators entertainment and DC |
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Beyond Washington Application Deadline |
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Washington DC Premiere of “An Almost Ordinary Summer” |
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La lógica perversa del sistema judicial estadounidense: de Guantánamo a Washington |
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Washington Column |
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NO CASE IS TOO BIG OR TOO SMALL FOR WASHINGTON LAW CENTER FIND OUT HOW WE CAN HELP YOUR CASE TODAY! |
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SBIA Government Relations Update: Washington Year in Review |
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Quer aprender inglês mas não sabe por onde começar Faça um tour 360º e veja porque nossos alunos indicam a Washington |
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Amusement Park in the Virginia amp Washington DC Area |
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DEPARTMENT OF STATE WASHINGTON 3d |
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The Washington Redskins Must Draft Quarterback Tua Tagovailoa |
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Location: The Reflecting Pool 1 Dr Washington DC 200001 |
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Conteo regresivo del Torito para su presentación en Washington Heights |
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Kimberly Hartz Named New CEO of Washington Hospital Healthcare System |
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Mr Tabith M Awal elder son of Mr Abdul Awal Mintoo and Mrs Nasreen Fatema Awal was born in a renowned Muslim family on 20th February 1979 He completed his education from George Washington University in DC USA and obtained combined degree in MSc in information System Technology and BBA in Management Information System |
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KALISPEL GOLF amp COUNTRY CLUB Spokane Washington |
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Oscar Nominations 2017: Denzel Washington Mahershala Ali Ruth Negga Octavia Spencer More |
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Washington Hospital CEO of 25 Years Nancy D Farber to Retire |
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Hard Money Loans for Seattle Tacoma Spokane and All of Washington |
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Coupon For Valentine’s Soul Jam Feat In Washington DC |
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10 Tiny Houses for Sale in Washington State |
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Mario Bemer USA Trunk Show in Washington DC |
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Embassy of the United Republic of Tanzania Washington USA |
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Weights from Toppenish Washington |
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Sr Director Digital Consumer Services Company Greater Washington DC area |
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Governor right to fight to open coal port in Washington state |
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Washington Monthly ranks RMU’s Graduation Rate 52 higher than expected based on its student population |
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12 Pioneering Facts about George Washington |
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Alabama Alaska Arizona Arkansas California Colorado Connecticut Delaware District of Columbia Florida Georgia Hawaii Idaho Illinois Indiana Iowa Kansas Kentucky Louisiana Maine Maryland Massachusetts Michigan Minnesota Mississippi Missouri Montana Nebraska Nevada New Hampshire New Jersey New Mexico New York North Carolina North Dakota Ohio Oklahoma Oregon Pennsylvania Rhode Island South Carolina South Dakota Tennessee Texas Utah Vermont Virginia Washington West Virginia Wisconsin Wyoming |
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Cyberattaque Wannacry: Washington accuse la Corée du Nord |
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2016 Washington Auto Show |
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Check out my new ‘Policy Watch’ for the Washington Institute: “Foreign Jihadists in Syria: Tracking Recruitment Networks” |
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Washington Wizards v San Antonio Spurs – NBA |
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Prep basketball roundup: George Washington boys sweep regular-season series from South Charleston |
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Washington Township resident says property littered with bullets from nearby gun club |
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George Washington At Yorktown Virginia |
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A journey up Mount Washington via Dog Mountain |
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Alex Washington DC USA |
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Washington Winegrowers |
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Washington Tehran step back from brink of war |
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Знал об угрозе Washington Post США |
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University of Washington President Emeritus Henry Schmitz drowns in Dungeness Bay on January 30 1965 |
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We are going to Brazil Washington – Salvador – Stopover in Cabo VerdeFebruary 20-25th |
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CJ Elleby’s 3-pointer leads Washington State to come-from behind victory over ASU |
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We are an advertising design and marketing agency headquartered in the greater Washington DC metropolitan area |
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Best iOS App Development Agency San Francisco California amp Washington |
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Marshall A Burke Joins the IRR-Washington DC Office as Managing Director |
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Unnatural History Amerika Cartoon Network kanalında yayınlanan gizem dolu bir gençlik dizisidir Henry Griffin antropolog olan annesi ve babasıyla tüm dünyayı gezmiş ve keşişlerden vücuduna hükmedebilmeyi öğrenmiş doğaüstü güçlere sahip bir gençtir Sahip olduğu bu doğaüstü güçler Henry’nin başına sık sık bela açmaya başlayınca annesi Jessica Henry’i Washington’da ki dayısının yanına göndermeye karar veriyor Dayısının yaşadığı kasabada çok sevdiği vaftiz babasının yaşadığını öğrenen Henry kasabaya taşındığında onun ölmüş olduğunu öğreniyor Kalp krizi sebebiyle öldüğü söylenen vaftiz babasının aslında cinayete kurban gittiğini düşünen Henry dayısının oğlu Jasper ile birlikte ipuçları armaya ve olayın gerçek yüzünü açığa çıkarmaya çalışıyorlar Her bölümde başka bir gizemin peşine düşen Henry Jasper ve arkadaşları Maggie’yi Türkiye’de Cnbce-e kanalından takip edebilirsiniz |
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Pre-Licensing and Continuing Education Classes in Maryland Washington DC amp Virginia |
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Trump’s impeachment defense waves away his chief of staff’s admission of quid pro quo – The Washington Post |
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KERRY WASHINGTON LOOKING BEAUTIFUL AT ABC’S SCANDAL ATAS Event-CAST ARRIVALS PHOTOS |
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Did you know the seventh-inning stretch originated in Washington DC In fact we have President William Howard Taft to thank for this tradition T’was on April 14 1910 as the Washington Senators played the Philadelphia Athletics at Griffith Stadium in Washington DC After sitting for a long period 6’2 300-pound President Taft stood up to stretch midway through the seventh-inning The entire crowd consequently rose to their feet as a show of respect On this day Taft also became the first US president to throw the first pitch |
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Back to Work! I have no idea whether I am on the tree or not However I received my PhD at the University of Washington in 1987 and wrote my dissertation under Paul Dietrichson whom I believe wrote his dissertation at Yale under either Brand Blanshard or Paul Weiss in the early 1950’s – Steven M Duncan I received my PhD at MIT in 1990 with George Boolos as my dissertation supervisor I currently teach philosophy at the University of Maine and would be pleased to be added to George’s tree -James Page Josh I’m very impressed at how you are talking to us from the future The last two posts have December 2006 datestamps! So here’s what I want to know:Who won the election Extra points for figuring out the temporal semantics and pragmatics of asking such questions about an event one month into the future of someone who lives two months in the future I think I can use either the past tense or the future tense– Kai Peter van Inwagen goes under Richard TaylorI assume there will be some interesting people whose dissertations he directed at Syracuse and ND but I’m not sure who exactly Dean Zimmerman It would be a good idea to add the year of receiving the PhDAnother issue: would you add Romanian philosophers as well Here is some information:Stefan Georgescu – Mircea Dumitru 1998 University of Bucharest School of PhilosophyGraeme Forbes – Mircea Dumitru 1998Mircea Flonta – Mihail-Radu Solcan 1993 University of Bucharest School of Philosophy – Adrian-Paul Iliescu 1983 University of Bucharest School of Philosophy – Gheorghe Stefanov 2001 University of Bucharest School of PhilosophyIlie Pirvu – Ioan-Lucian Muntean 2005 University of Bucharest School of PhilosophyIon Ianosi – Gabriel Liiceanu 1976 University of Bucharest School of Philosophy – Catalin Cioaba 2005 University of Bucharest School of Philosophy – Sorin Lavric 2005 University of Bucharest School of PhilosophyAlain de Libera – Stefan Vianu 2001 University of Geneva Just thought I would add someone JM Bernstein Chair at The New School for Social Research who recieved his PhD from WH Walsh at the University of Edinburgh in 1975 I finished my DPhil in Oxford in 1962 on Knowing and Understanding — an attempt to defend Kant’s claim that mathematical knowledge was both synthetic substantial non-empirical and necessary My first degree was in mathematics and physics and most of what I heard philosophers say around that time about the the nature of mathematical knowledge was clearly wrongMy official supervisor was David Pears but I saw little of him and learnt much more from my ‘personal tutor’ at Balliol College Richard Hare and people whose seminars I attended eg Gilbert Ryle Michael Dummett John Austin Elizabeth Anscombe Geoffrey Warnock and fellow students From about 1970 I started to realise that the best way to do philosophy is to combine it with AI That’s also the best way to do AIAaron Sloman What a great idea! I noticed my advisor’s name on here and was delighted to find that my lineage traced back through Leo Strauss and Ernst Cassirer to Kant and Leibniz I suddenly feel very dignified and inclined to add my name to the tree if you’re still updating I received my PhD in 2000 at Penn State with Dan Conway as my dissertation adviser-Pete Groff C I Lewis supervised WV Quine who supervised William CraigJohn Myhill Hao Wang Hughes Leblanc and Robert McNaughton who supervised me John Corcoran–already immortalized on this blogI supervised George Weaver who supervised Sean GallagherI also supervised Michael Maloney Edward Keenan Terry Nutter Susan Wood Michael Scanlan and Sriram Nambiar-John Corcoran Some Terrence Irwin: Susan Meyer is the same person as Susan Saude the former is her married name Under Rawls Onora O’neil is the same person as Onora Nell former is her married name Alyssa Bernstein is the student of Bonnie Steinbeck student of Vermazen student of DavidsonMatthew Katz is the student of Scott Weinstein student of Saul KripkeMark Navin is the student of Kok-Chor Tan student of Frank Cunningham student of David Gauthier Note that Tan and so Navin should also be listed as a student of David Dyzenhaus co-adviserScott Edgar Marc Cohen and Morgan Wallhagen should be listed as students of Gary Hatfield student of Dretske student of Broadbeck student of BergmannKate Moran and Jennifer Dobe should be listed as students of Paul Guyer student of Cavell student of AustinCarol Hay should be listed as a student of Louise Antony student I think of PutnamElizabeth Herschbach and Todd Bates should be listed as students of James RossAccording to the ‘memorial notice’ by Kathleen Higgins Robert Solomon should be listed as a student of Frithojoff Bergmann Just trying to fill in my own tree someI studied under James Lindemann Nelson already in tree as child of Richard HullOther advisees include Maya Goldenberg Lisa Eckenwiler Matthew Stolick Christopher Crittenden Carolyn Ells- Rory E Kraft Jr Elia Zardini currently at Univ of St Andrews was advised by Crispin WrightEline Busck Gundersen currently at Univ of Oslo was advised by Crispin WrightIwao Hirose currently at McGill Univ was advised by John SkorupskiEnzo Rossi currently at Univ of Wales Newport was advised by John SkorupskiRaffaele Rodogno currently at Swiss National Science Foundation was advised by John SkorupskiKent Hurtig Currently at Univ of Stirling was advised by John Skorupski Thomas Reid: C Stove: I’m not sure if he’s still listed as an orphan or not he was for a long time but if so Barry Stroud should be listed as a student of Morton White according to Stroud’s Dewey lectures in the most recent APA proceedings White was his dissertation director at Harvard Is this still alive Would be a shame if not!! 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by Linda Wetzel Peter Kivy was the advisor of Gregg Horowitz VanderbiltTuebingen is where Manfred Frank teaches it is not his last name Thanks for all the updates and corrections and for pointing out the very embarrassing Tuebingen error — I think I must have been asleep when I entered that They’ve all been entered and will show up when I post the next update Rosemarie Tong is incorrectly listed as the student of Monroe Beardsley Her dissertation advisor was actually Elizabeth Lane BeardsleyAdditional students of Monroe Beardsley include: Michael J Wreen and Donald CallenAdditional students of Joseph Margolis include: Kathleen J Wininger James Stieb James Crippen Jr Rosalind Hursthouse Auckland was advised by GEM Anscombe On page 90 of the list Rebecca Comay is listed as an orphan She was supervised by Graeme Nicholson who appears further down on the list in the George Sydney Brett thread Ausonio Marras an orphan on p 97 was advised by Romane Clark when he was at Duke Marc Ereshefsky an orphan was advised by Elliott Sober Ereshefsky has advised Jesse Hendrikse2005 Jay Odenbaugh 2001 Baumslag 1997 and Mohamed El Samahi Michael Liston was co-advised at UC San Diego by Mark Wilson child of Hilary Putnam and Zeno Vendler child of Paul Ziff Dummett was my supervisorBrendan Frank Cunningham an orphan on p 90 was supervised by David Gauthier graduating in 1970 when Gauthier was at Toronto Joseph Cohen was supervised by Jacques Derrida and was apparently Derrida’s last studentGregory Recco was supervised by John Russon and graduated in 2002 John Russon was supervised by Graeme Nicholson graduating in 1990 Russon supervised David Ciavatta Nathan Anderson Eckerd College was advised by John Russon Completed in 2002 Peter Costello Providence College was advised by John Russon Completed 2002 W Bruce Gilbert Bishop’s University was advised by John Russon Completed 2002 Brian Mackintosh University of Alabama was advised by John Russon Completed 2000 Michael Svoboda George Washington University was advised by John Russon Completed 2002 Kym Maclaren Ryerson University was advised by John Russon Completed 2004 Kirsten Jacobson University of Maine was advised by John RussonCompleted 2006 Maria Talero University of Colorado at Denver was advised by John Sallis Completed 2002 Eric Sanday Marquette University was advised by John Sallis Completed 2002 Robert Metcalf University of Colorado at Denver was advised by John Sallis Jay Lampert University of Guelph was advised by Henry Pietersma at the University of Toronto Completed 1987 James Vernon York University was advised by Jay Lampert at University of Guelph Antonio Calcagno Scranton was advised by Jay Lampert David Morris Trent University was advised by Graeme Nicholson James Crooks Bishop’s University was advised by Graeme Nicholson Bernard Freydberg Slippery Rock University was advised by John C Sallis Claudia Baracchi New School for Social Research was supervised by John C Sallis Daniel Shannon DePauw University was advised by Henry Harris Tamar Japaridze Tblisi was advised by Rebecca Comay Andrea Sauder was advised by Rebecca Comay Emilia Angelova Trent University was advised by Rebecca Comay Ingo Brigandt University of Alberta was advised by Paul Griffiths and Anil Gupta Gordon Bowie is actually Lee Bowie I believe the last time the name Gordon was used was on his my dissertation 35 years ago For many years he was known as G Lee Either G Lee Bowie or Lee Bowie would be fine solutionsLee Bowie Omer Aygun was supervised by John Russon completed in 2006 I discovered 3 errors on the list: 1 Jerrrold Katz wrote his dissertation on the impossibility of justifyinginduction with me not with Alonzo Church I don’t think Church had anything to do with it 2 I directed Raymond Smulyan’s dissertation and not Church although Church signed off on it as well since I was not at that point a member of the Mathematics DepartmentI should explain that at Princeton MIT and Harvard I directedPhD work in the mathematics department as well as in the philosophydepartment In fact In 1960-61 I was made a formal member of the Princeton Mathematics Department and Church was made a formal member of the Philosophy Dept so that each of us had tenure in both departments Also 3 Harold Hodes wrote his dissertation under my supervision although it was an MIT dissertation and I was already at Harvard and we published the main results as a joint paper: An Intrinsic Characterization of the Hierarchy of Constructible Sets of Integers Stephen Leeds and Hilary Putnam Logic Colloquium ’69 eds R O Grandy and C E M Yates Amsterdam: North-Holland 1971 pp 311-350 Although James Thomson whom you list as directing it signed off on the dissertation I doubt that he had the mathematical knowledge to even follow it let alone direct it Probably you are not interested in my PhD students inmathematical logic unless they received philosophy degrees as didRIchard Boyd and George Boolos whom you list but here they are:At Princeton there were two: Raymond Smullyan and Robert W Ritchie At MIT in addition to George Boolos whom you have correctly listed and Harold Hodes see above there were Stephen L Bloom Gustav Hensel C Ward Henson David Isles David Luckham Hartley Rogers signed off on the thesis Miriam Lucian and Joan LukasAt Harvard there was Herb Enderton Also my wife Ruth Anna Putnam is listed as Carnap’s studentwhich she was but under her name from her first marriage RuthMathers which she has never published under Since she is well knownit would be good if this can be changedThe tree is wonderful! I only wish there were some way to cause aname to light up in its location on the tree Sincerely yours and thanks for the tree Hilary PutnamCogan University Professor Emeritus Harvard University Christian Wuthrich now at the University of California San Diego was a student of John Earman Pittsburgh RG Frey now listed as an orphan should be listed as a son of R M Hare John Cooper student of Gwil Owen was the advisor for Cass Weller 1983 Cass Weller was the advisor for Audre Brokes 1997 Cass Weller was the advisor for David Nixon 2004 Cass Weller was the advisor for Ben Stenberg 2006 John Zeis Canisius College Buffalo did his dissertation with Geach Anscombe and James Ross at U Penn Andrew Cullison West Virginia did his dissertation with Richard Feldman Trendelenburg supervised Jurgen Bona Meyer It looks like he also supervised Friedrich Paulsen according to Klaus Kohnke’s book The Rise of Neo-Kantianism I was wrong about Paulsen’s advisor Kohnke meant just that Trendelenburg supervised Paulsen’s undergrad dissertation 1871 According to Paulsen’s Autobiography Columbia Univ Press 1938 his Habilitation allowing him to teach as a privatdozent was passed by Friedrich Harms and Eduard Zeller 1875 Zeller seems to have been the more influential examiner Alois Riehl supervised Adhemar Gelb at Berlin Gelb a psychologist graduate c 1910 Elizabeth Barnes was advised by Daniel NolanMarcus Rossberg was advised by Crispin WrightEline Busck Gundersen was advised by Crispin WrightMarkus Schlosser was advised by Sarah BroadieMichael Weh was advised by Berys Gaut Jon Williamson Kent currently was advised by Donald Gillies Harold Hodes advised Jeffrey Roland Cornell and Mirek Janusz also Cornell Nicholas Rescher has an article in Erkenntnis tracing the development of a Berlin school Info there inc the following:Laurence Laudan was supervised by C C GillispieRescher says that Adolf Grunbaum’s dissertation was directed by Hempel at Yale 1950 not ReichenbachRescher also says that Reichenbach supervised most of Hempel’s work but that he left before Hempel’s dissertation was approved The psychologist Wolfgang Kohler stepped in to complete the formalities of the degree says RescherLaura Ruetsche and Gordon Belot were supervised by John EarmanBruce Buchanan was supervised by Gerald Massey David Hershenov was advised by Nathan Salmon in 2002 Hi just a correction You have two listings in different places on the tree Earline Ashworth and Jennifer Ashworth These are both the same person Most of her publications are under the name EJ Ashworth just to throw a third name into the mix She was my supervisor at UW David Wong currently at Duke was a student of Gil Harman Some more students of Kazimierz Twardowski:-Roman Ingarden-Tadeusz Czeżowski-Jan Łukasiewicz-Izydora Dąmbska-Władysław Witwicki David L Hildebrand Asst Professor Univ of CO at Denver advised by Douglas Browning UT Austin Ingo Brigandt now at the University of Alberta was a student of Paul Griffiths CI Lewis was the advisor of Kenneth SayreI believe that Farber should be listed as Sellars’s advisor My grounds for saying this are that I once circa 1982 or 83 had a conversation with Sellars about Husserl and he pointed out that Husserl was his teacher’s teacher identifying Farber as the intermediate linkThe Farber-Husserl link is also difficult however as Farber did his PhD at Harvard though Husserl was almost undoubtedly his most significant philosophical influence and teacher This isn’t very important but when he gave a talk here at Cornell in the fall Ralph Wedgwood happened to mention that his primary adviser wasn’t in fact Nick Sturgeon as currently listed on the tree but Terry Irwin Roderick Firth was Clement J Dore’s advisor Melissa Lane PhD in Philosophy from Cambridge now employed in teaching history of philosophy and political thought in Cambridge History Faculty was advised by Myles Burnyeat and Malcolm Schofield Hans Reichenbach’s dissertation advisor was not EMMY Noether as mentioned in the database but her father MAX Noether Since Reichenbach has quite a few descendants it’s somewhat surprising that nobody has noticed this before I assume you’ve takenthe information from the Mathematics Genealogy Database which alsohad the wrong information but they have corrected it now Some miscellanea:Jocelyn Dunphy-Blomfield Monash was supervised by Paul Ricoeur Sorbonnewho was taught by Roland Dalbiez RennesHannah Arendt New School for Social Research was supervised by Karl Jaspers Heidelberg who was supervised by Edmund Husserl Freiburg im Breisgau who was supervised by Franz Brentano Vienna Ágnes Heller New School of Social Research was supervised by György Lukács Budapest Two recent students of John Broome Oxford:James Morauta – now at University College OxfordAndrew Reisner – now at McGill There are a few more advisees of Ron Chrisley who now have their doctorates: Tony Morse Marek McGann Marzieh Asgari-Targhi and Hanne de Jaegher John Rawls was main advisor and TM Scanlon second advisor both Harvard University for Andreas Follesdal University of Oslowho was main advisor for Jacob Elster — continuing above comment:Andreas Follesdal has advisedJakob ElsterLars Fjell HanssonOle Frithjof Norheim Chris Campolo was advised by Georgia Warnke Mikkel Gerken was advised by Tyler BurgeThanks for the tree! You are doing the philosophical community a real service Best regardsMikkel Gerken Gillian Barker was advised by Philip Kitcher Bradley Wilson Slippery Rock University was advised by William Lycan Robert Bass was advised by Edward F McClennen Nathan Hanna Syracuse was advised by Edward F McClennen and Kenneth Baynes RE: G A Press Speaking as himself I would add this Jason Saunders was the director of my Ph D dissertation Other important teachers were: Richard H Popkin Paul Henry SJ and Herbert Marcuse As a specialist in the history of ancient philosophy my work reflects the orientations of several of my teachers’ teachers: Paul O Kristeller Frederick J E Woodbridge and John Herman Randall Jr My orientation was also influenced early on my Richard P McKeon RE: Richard H Popkin although Nagel was indeed his dissertation director his own scholarship owed more to Paul O Kristeller and John Herman Randall R T McIntyre would prefer to be listed as Ronald McIntyre Dagfinn Follesdal along with Jaakko Hintikka was co-advisor Follesdal was major influence on subsequent work Manfred Frank was advised by Dieter Henrich in fact Henrich was his co-advisor but he is by far the more important intellectual influence than his actual advisor who was not even a philosopher but a professor in German literature among his advisees are Thomas Grundmann who is the advisor of Magdalena Balcerak Frank Hofmann Catrin Misselhorn Gianfranco Soldati Doris Gerber According to his obituary in the Proceedings and Addresses of the American Philosophical Association 1970 Albert Hammond was AO Lovejoy’s student And I believe Norman Kretzmann was Albert Hammond’s student Finally Scott MacDonald was Norman Kretzmann’s student That should incorporate a few orphans into the already sizable Leibniz tree Stephanie Bauer Washington University in St Louis May 2008 was advised by Marilyn Friedman Hi JoshNeat projectI am already on the Tree but my dissertation was co-advised so rather than just Robert Barnard student of John Tienson it should also include student of Terry Horgan — this would create interesting issues in real geneticsAn additional item: I was looking up my colleague Neil A Manson student of Peter Van Inwagen and found that your index currently lists only his professional doppleganger Neil C Manson ThanksBob Barnard Dylan Dodd PhD UC Santa Barbara 2006 was a student of Anthony Brueckner Benjamin Huff completed a PhD in 2006 under advisor David K O’Connor who also advised Dan Moloney and Andrew PayneTwo corrections: David Solomon advised Angela Faulconer a is missing in the PDF and I suspect in 1985 he advised only one Mark Theodore Nelson who seems to appear both at the start and finish of his list Peter Simons Leeds advised Sarah Kattau 1999 Sandra Lapointe 2000 and Bo Meinertsen 2005 RE: Dieter Henrich as supervisor of Raymond Geuss Henrich was indeed on Geuss’ dissertation committee and a continuing presence at Columbia in the late 1960s and early 70s when Geuss was a graduate student there But the main influences on Geuss in philosophy at that time were his dissertation supervisor Robert Denoon Cumming and Sidney Morgenbesser Caery Evangelist finished her PhD with Edward P Mahoney Duke 2004 Eric Steinhart was advised by Eva Kittay T K Seung University of Texas at Austin advised the dissertations of Michael Locke McLendon Cal State–Los Angeles Sharon K Vaughan Morehouse College and David Lay Williams University of Wisconsin–Stevens Point Gordon Barnes was advised by Alan Sidelle Robert Nozick advised Steven Luper Teddy Seidenfeld was advised by Isaac Levi Jeffrey Helzner was advised by Seidenfeld Ignore the last comment as both Helzner and Seidenfeld are already on the tree Matt Lister was advised by Samuel Freeman who was advised by John Rawls Ryan Muldoon was advised by Cristina Bicchieri I’m not sure if Bicchieri is on the tree yet or not She did her PhD in the history and philosophy of science department at Cambridge where she was a student of Mary Hesse Cristina Bicchieri also advised Giacomo Sillari Josh will you please add Matthew Van Cleave 2007 and Giovanni Mion 2008 as students of mine And if you want to add a year to mine it was 1984 Thank you Chris Gauker A correction: Ronald de Sousa Univ of Toronto is listed twice once under ‘DeSousa’ and once under ‘Sousa de’ Under one of these names he’s an orphan Elizabeth Schier Macquaire University PhD Adelaide University 2007 was supervised by Gerard O’Brien and Jon Opie This would be a thread going out from Trendelenburg but it is not complete at all:Marcela Garcia daughter ofThomas Buchheim son ofRobert Spaemann son ofJoachim von Ritter son ofErnst Cassirer son of Paul Natorp son ofErnst Laas son ofF A Trendelenburg walter j schultz should be added to the tree Who knows where to download XRumer 50 Palladium Help please All recommend this program to effectively advertise on the Internet this is the best program! Correction: Marty Heitz was student of Graham Parkes not Graham PerkinsAddition: Angela Potochnik was student of Peter Godfrey-Smith John A Baker was my PhD supervisor at the University of Calgary where I graduated in 1980 A list of all of John’s students can be found PfeiferUniversity of Saskatchewan Michel Foucault’s supervisor: Georges Canguilhem Alfred Louch’s advisor was John Wisdom Louch is orphaned on the list Justus Buchler’s dissertation director was Ernest Nagel This still places him in the Leibniz line but through Royce not Dewey Buchler also studied with Dewey Deborah Achtenberg was advised by Stewart Umphrey Albert Hofstadter 1910–1989 was advised by Frederick J E Woodbridge 1867-1940 at Columbia Mary Mothersill Columbia deceasedwas advised by CI Lewis Craig Vasey was advised by Paul Ricoeur at Universite de Paris X for his Doctorat du troisieme cycle 1982 Also by Emmanuel Levinas at Universite de Paris IV for his Maitrise 1978 The current information Roderick Chisholm Brown U is also correct — PhD 1984 Craig Vasey was advised by Paul Ricoeur at Universite de Paris X for his Doctorat du troisieme cycle 1982 Also by Emmanuel Levinas at Universite de Paris IV for his Maitrise 1978 The current information Roderick Chisholm Brown U is also correct — PhD 1984duversin Keith Frankish was advised by Peter Carruthers Myles Burnyeat was supervised by Bernard WilliamsAdditional students of Myles Burnyeat include:Angela HobbsAdditional students of Peter Strawson inclue:Quassim Cassam Katherine Hawley St Andrews goes back via Peter Lipton Cambridge to AJ Ayer Lei Zhong Michigan 2010 was advised by Peter Railton what a great blogdissertation help I would like to thank you for the efforts you have made in writing this article Common digestive system diseases and disorders Health benefits of saltHealth benefits of salt Prevent wrinkles naturally Hemorrhoids Toothache pain home remedies Hip pain Top fertility foods Dandruff treatment Garlic benefitsHealth benefits of garlic Remedies for sinus headache Get smooth and glowing skinSmooth and glowing skin Cellulite Tighten sagging skin Dry skin patches treatmentDry skin patches Get rid of acneGet rid of acne Driving tips Migraine headache remedies Migraine headache Sue Larson PhD Standford taught at Barnard/Columbia was advised by Donald Davidson and Sue was my advisor My PhD is from Columbia and I teach at Mills College More effort should be made to throw more light on this topic very interesting thanks:- xxx Click here for Student Escorts London OSHA 10 hour Explain the OSHA 10 hour course training Took a lot of time to read but I really found this very interesting and informative thank you buddy for sharingCustom Logo DesignStationary Design ServicesWebsite Design ServicesBanner Design ServicesBrochure Design Services Apple unveiled a new update to the iPod Nano today showing off a few new features that will excite fitness nuts and watch loverswrist watches for menmens wrist watchesmens sports watchesmens digital watcheshigh quality watchesdesigner mens watches Took a lot of time to read but I really found this very interesting and informative thank you buddy for sharing I hope someone updates the tree to make corrections and additions as in these comments I have been searching for such information for months now Thanks God I have found your blog it has helped me a lot in working on my project Thanks a lotlogo design price Your post is awesome You have explained it very well I am willing to read posts like this one Thanks for postingDissertation Writing Help It is an excellent post You have written it very well It has helped me a lot in my research Keep on posting such articles I will tell my friends about your website Thank youwriting custom term papers What an enjoyable read Cell Phone Lookup this turned out to be! 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John Hawthorne took his doctorate from the department in this period if I’m not mistaken as did Mark Heller That forms a pretty interesting MampE tree Hawthorne returned to Syracuse to teach in the late 1990s and was later joined by Ted Sider from Rochester and then Dean Zimmerman from Notre Dame This was after Bennett retired and Van Inwagen left for Notre Dame himself but before the three young metaphysicians were collectively recruited to the Rutgers department in 2002 dealing a major loss to Syracuse I’m not sure if such a singular blow is unprecedented or not But it was huge and controversial even within the Rutgers department Brian Weatherson left the department around the same timeThe Syracuse department has throughout maintained its strength in moral philosophy and especially moral psychology Michael Stocker Laurence Thomas and Lynn McFall now emeritus all taught in the field throughout the 1990sThe historical strength in modern philosophy which was provided by Jonathan Bennett eventually led to the hiring of Fredrick Beiser who is known for his work on German idealism and the English Enlightenment Kenneth Baynes has been a recent hire whose interests in modern and contemporary German thought compliment and create a new strength for the departmentDespite the losses to Rutgers Syracuse still maintains significant strength as a center for the study of the metaphysics of identity and the subject of persistence through time Andre Gallois and Mark Heller replaced Ted Sider Kris McDaniel is a junior faculty hire with interests in the field There is a nice history of the Philosophy Department at Washington University in St Louis to be found with a detailed history of the department itself it covers the antecedents of the department as wellIt was researched and written by Richard Red Watson now Emeritus at Wash U best best best best best site site site best Blogs are so informative where we get lots of information on any topic Nice job keep it up!!Dissertation 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Carnegie Institution of Washington Publication Volume 248 2 248 2 |
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Patricia C Wright Vololontiana R Razafindratsita Sharon T PochroN Jukka Jernvall The Key to Madagascar Frugivores Abstract In the Malagasy ecosystem one particular animal group lemurs have the greatest biomass and species richness of frugivores The peak fruit production in the Malagasy rain forests is about three months shorter compared to peak fruit production in the Amazon and the African rain forests This suggests that the environment in Madagascar has more well-defined constraints than other continental areas with primates In Ranomafana National Park both the overall number of trees and the number of tree species producing fruit drops during winter months Particularly in large-bodied lemurs such as sifakas drop in fruit availability corresponds to an increase in leaf eating In addition to dietary shifts all lemur species appear to be able to deal with the season of scarce fruit availability by conserving energy Extreme responses to winter season are seen in small-bodied lemurs which go into hibernation up to six months every year Unlike many primate communities in other continents lemurs do not have synchronous birth peaks across species In Ranomafana sympatric lemurs show that while individuals within a species have synchronized births across species lemurs have synchronized weaning The weaning synchrony coincides with maximum fruit availability and production of small fruits peaks when small juvenile lemurs begin to forage independently These patterns suggest that lemurs do not appear to rely on fruits to carry them over the period of food scarcity as would be expected from classical descriptions of keystone resources Rather we propose that lemurs as a guild rely on fruits as a keystone resource during the warm wet months in order for lactation and weaning to succeed Many of the fruit tree species used by lemurs are also hardwood species favored by selective loggers While loss of these key fruit trees may not drive lemurs into extinction immediately it may adversely affect reproductive success years after logging Key Words: Frugivory weaning synchrony color vision Ranomafana National Park Madagascar INTRODUCTION Madagascar the fourth largest island in the world rifted from Africa more than 150 million years ago and has been isolated in its present position for over 88 million years Krause 1997 Over 1500 km long Madagascar supports as rich and varied a flora as can be found anywhere in the tropics Schatz et al 2002 including rain forest dry subtropical forest and spiny desert Botanists surveying 3 one ha plots in the Ranomafana National Park RNP rain forests have found 37 families and 105 species of trees Schatz and Malcomber in Wright 1997a These RNP plots were not as diverse as one ha plots in Colombia with 44 families and 197 species but were still more diverse than an African lowland forest in Gabon 29 families and 99 species Gentry 1993 Reitsma 1988 Many animal taxa are absent from Madagascar because of biogeographical history and ungulates monkeys many birds and many bats simply never reached the island What effect does this absence have on the ecology of the rain forest In most rain forests the primary pollinators and seed dispersers are insects birds and bats Bawa et al 1990 Fleming et al 1987 Terborgh 1986 estimated that as much as 80 of Amazonia’s mammalian biomass depends on fruit resources As pointed out by Fleming et al 1987 in contrast in Madagascar 68 of the birds are insectivores and only 8 are frugivores The only avian seed dispersers in the rain forest are the velvet asity Philepitta castanea in the understory and the Madagascar bulbul Hypsipetes madagascariensis in the canopy Langrand 1990 Razafindratsita 1995 Four bird species are nectivores and pollinators Two species of parrots and several species of pigeons are seed predators Bats are also relatively depauperate in diversity with 28 species and almost all are insectivorous Peterson et al 1995 Lemurs may be the primary pollinators and seed dispersers in the rain forests of Madagascar Balko 1998 Kress et al 1992 Nilsson et al 1993 Overdorff 1992 Ratsimbazafy 2002 Wright amp Martin 1995 Black and white ruffed lemurs red-bellied lemurs and brown lemurs medium sized 2-4kg diurnal diurnal primates pass vine and tree seeds intact and these sprout faster and with less mortality than seeds not passed through a primate gut Dew amp Wright 1998 Indeed in the Malagasy ecosystem it is actually possible to single out one particular animal group lemurs which have the greatest biomass of frugivores and may qualify as keystone mutualists Gilbert 1980 Almost all lemur taxa eat some fruit but despite this some fruits such as figs are much rarer in Madagascar than in other tropical forests where they are described as keystone food sources for monkeys Goodman amp Ganzhorn 1997 Terborgh 1983 Although in other tropical rain forests such as Kibale Forest see Chapman et al this volume and BCI see Milton et al this volume long-term phenology data show that fruit production varies greatly for individual trees there are fruits available in those forests all year Struhsaker 1997 And in rain forests such as Manu Park in the Peruvian Amazon keystone resources such as figs large-crowned or nectar abundant patches provide food for frugivores during extended periods of fruit scarcity Terborgh 1983 Wright 1989 Madagascar rain forests unlike other forests with 12-14 species of sympatric primates have a much longer period without fruits up to six months a year Wright 1999 These long periods without fruits are reflected in the fact that few lemurs are obligate frugivores Dew amp Wright 1998 Fleming et al 1987 Goodman amp Ganzhorn 1997 Overdorff amp Strait 1998 Tattersall 1982 but nonetheless fruits seeds and flowers compose 40-90 of the annual diet of Eulemur Varecia Eulemur ssp Propithecus Microcebus Cheirogaleus and perhaps Mirza and Phaner Atsalis 1999 Balko 1998 Ganzhorn amp Kappeler 1996 Ganzhorn et al 1999 Hemingway 1996 1998 Overdorff 1991 1993 Overdorff and Strait 1998 Powzyk 1997 Wright amp Martin 1995 In this paper we examine fruiting patterns in the rain forest of Madagascar paying particular attention to the degree of seasonality of fruit production in relation to lemur life history To understand long term patterns in detail we examine feeding and reproduction in the largest species of lemur 6kg the Milne Edwards sifaka Propithecus diadema edwardsi The feeding behavior of this species is compared with fruit availability in the forest using fruiting phenology of 98 plant species Finally we compare fruiting with key life history events among several sympatric lemurs We propose that for lemurs fruits are keystone foods critical for lactation and reproductive success rather than for survival during harsh years METHODS Study Site Ranomafana National Park established in 1991 is 43500 ha of continuous rain forest located in southeastern Madagascar at 21 16’S latitude and 47 20’E longitude Wright 1992 Wright and Andriamihaja 2002 The park is 25 km from Fianarantsoa and 60 km from the Indian Ocean Elevations range from 500-1500m within the park and annual rainfall ranges from 1600-4017 mm RNP records Most of the rain falls during the months from December to March Temperatures range from lows in June-September 4-12C to highs in December-February from 30-32C The study groups of sifakas were located in the 5km2 Talatakely study site TTS which was selectively logged by hand in 1986-1989 The park contains moist evergreen forest and the canopy height ranges from 18m-25m Phenology data have also been taken at an unexploited site within the continuous forest of the park 3km south of Talatakely at Vatoharanana Hemingway 1996 Overdorff 1991 and a third unexploited site 3km further South at Valohoaka Balko 1998 Botanically RNP is one of the most diverse rain forests in the world Lowry et al 1997 This area has had a non-hunting tradition and impact of human predation on lemurs and viverrids has been minimal over at least the last 50 years Wright 1997a The faunal diversity in Ranomafana National Park RNP is high for Madagascar Wright 1992 1997a with 114 species of birds including six species of raptors six species of viverrid and twelve species of primates Table 1 Wright 1998 Razafindratsita 1995 Sifaka biomass estimates were 125kg/km2 Wright 1998 Total biomass of primates at this site was 330kg/km2 comparable to terra firme forests in Central Amazon and Lope Reserve in Gabon but roughly half the primate biomass of the alluvial flood plain forest of Manu Peru or Kirindy dry forest in western Madagascar Ganzhorn amp Kappeler 1996 Oates et al 1990 Peres 1993 Terborgh 1983 Wright 1998 Table 1 Lemur species in the rainforest site of Ranomafana Lemurs with an annual diet of at least 30 fruits are marked with asterisk Only one diurnal species Varecia variegata eats fruits for over 85 of its diet Balko 1998 Ratsimbazafy 2002 Species Body Mass g Biomass kg/km2 Avahi laniger woolly lemur 900 18 Propithecus edwardsi Milne Edward’s sifaka 5800 125 Cheirogaleus major fat-tailed dwarf lemur 350 18 Microcebus rufus rufous mouse lemur 42 4 Daubentonia madagascarienis aye-aye 3500 7 Lepilemur microdon sportive lemur 970 16 Hapalemur griseus grey gentle lemur 935 20 Hapalemur aureus golden bamboo lemur 1550 96 Hapalemur simus greater bamboo lemur 2450 12 Eulemur fulvus rufus brown lemur 2200 66 Eulemur rubriventer red bellied lemur 2000 48 Varecia variegata variegata ruffed lemur 3650 9 Fruit Collection and Morphology During the 27 month time period April 1997-June 1999 fruits were collected in the forest by TR Paul Rasabo ICTE botanists and other research assistants All fleshy fruits from canopy trees understory trees bushes and vines were chosen regardless of the animal consumer in order to obtain a quantitative measure of fruit availability The parent plant was measured as to height dbh crown diameter crown depth and located on a map In the lab 10 fruits of each plant were weighed on a digital scale measured with calipers Color was noted Then the fleshy aril was removed and seeds were counted weighed and measured For analysis the fruits were grouped according to weight The small fruit category consisted of fruits that weighed 1g In addition fruits were grouped into four color categories « Dark » were black purple and brown while the category « red » were red and orange and « light » were white and yellow The fourth category was « green » which were green colored ripe fruits thus excluding green but unripe fruits Monitoring Fruit Phenology One to three individuals of 98 species a total of 233 trees of fruiting trees were chosen Morphological information including height dbh crown diameter crown depth were taken on each selected tree bush or vine The location of each study tree was mapped on the Talatakely trail map Twice each month from March 1997-February 1999 a botanist and several assistants observed each tree using binoculars to determine if a tree had fruits ripe and unripe flowers or new leaves A score of 1-5 was given on abundance of each category with five being high abundance Sifaka Fruit Feeding Patterns Within the same trail system as the phenology trees we have been conducting studies on the behavior and ecology of four adjacent groups of Milne Edward’s sifakas Propithecus diadema edwardsi since 1986 Pochron et al 2004 Pochron amp Wright 2002 Wright 1995 1998 1999 Individuals from these groups were captured with anesthetic darts weighed measured and given colored nylon collars for field recognition Glander et al 1992 Wright 1995 The demography and changes in group composition are described in detail by Pochron amp Wright 2004 and Wright 1995 For this paper we examined the proportion of feeding time spent feeding on fruits One individual sifaka per day was observed from awakening until asleep in the sleep tree for 2 days each month from January 1998-December 1998 During this time all foods eaten were identified and the number of five-minute samples during which each individual fed was recorded RESULTS AND DISCUSSION Seasonality in Fruiting and Fruit Feeding Despite the similarity of Madagascar in number of species and number of stems found in tropical forests of other geographic areas Abraham et al 1996 Lowry et al 1997 Sussman amp Rakotozafy 1994 the fruiting patterns in Madagascar contrast with many rain forests Ganzhorn et al 1999 Wright 1999 Madagascar phenology plots including the data shown in this paper show a prolonged season of 4-6 months with few trees in fruit Hemingway 1995 Overdorff 1993 Powzyk 1997 Sauther 1991 In addition in Malagasy rain forest many canopy species produce flowers and fruit on prolonged irregular asynchronous or alternate year cycles Hemingway 1995 Morland 1991 1993ab Overdorff 1993 Powzyk 1997 Figure 1A shows that of the 98 species studied the number of species that carry fruit cycles between 10 and 50 During the austral winter June-September of 1997 and 1998 the number of fruiting species remained below 25 and 20 respectively In contrast the times of abundant fruiting of individual trees scores 3 to 5 are typically during austral summer Fig 1A Thus Malagasy winters are not only characterized by a drop in the number of trees producing fruit but also a lack of species producing fruits Figure 1 Phenology of fruit availability in 98 species A and a subset of 31 species eaten by sifakas B in the rain forest of Ranomafana National Park during April 1997 to June 1999 Note how changes in number of fruit species eaten by sifakas track the overall richness of fruiting taxa Fruit score one represents even one fruit observed in a tree fruit score two represents up to a quarter of the branches having fruit and score five describes branches having abundant ripe fruits Months in parenthesis lack data Compared to Madagascar where fruit availability is highly seasonal fruiting occurs throughout the year in other tropical forests such as Kibale Forest Uganda Chapman et al this volume Struhsaker 1997 Ketambe Sumatra van Schaik 1986 Manu Peru Gentry amp Terborgh 1990 Terborgh 1983 Gabon Gautier-Hion et al 1985 Barro Colorado Island Panama Foster 1982 Milton et al this volume Colombia Stevenson this volume Manaus Brazil Lovejoy amp Bierregaard 1990 and Maraca Brazil Nunes 1998 with a tendency for slightly lower fruit production in the driest months usually for a two or three month period The seasonal cycling of Malagasy fruit diversity Fig 1A is strongly reflected in the diet of Propithecus diadema edwardsi During 1998 these sifakas ate fruits of 31 species out of the studied 98 Fruiting of the 31 species resembled closely the overall seasonal patterns Fig 1B However sifakas appear to increase the relative diversity of fruit species consumed during the summer season compared to winter Roughly 20 of concurrently fruiting species were eaten during the winter while up to 30 were eaten during the summer This suggests that sifakas are opportunistic frugivores that track the overall fruit richness in the forest This is further evident when the amount of fruit feeding is examined In Figure 2 the percentage of fruit feeding minutes is plotted for 1998 Sifakas spend about half of their feeding time on fruits during the summer while the proportion of fruit feeding drops all the way to zero in July Fig 2 There is a high turnover of top ranking fruits in the diet among months from the families Clusiaceae Myrtaceae and Lauraceae implicating again the opportunistic nature of fruit eating in sifakas The majority of sifakas’ feeding time is spent eating leaves Figure 2 Distribution of fruit feeding of total feeding time of sifakas during January 1998 to December 1998 and phenology of fruit availability Because less than half of the 98 sampled plant species carried fruit in any single month Fig 1A we also examined the overall turnover of fruiting species by calculating cumulative fruiting curves Fig 3 Beginning from a fruiting peak when roughly 42 of species carry fruit in 12 months over 70 of the species have fruited and in 24 months up to 85 of species have fruited Fig 3 These numbers suggest a relatively high turnover rate of fruiting species which suggests that the flexible fruit feeding of sifakas may be a good strategy in a Malagasy forest where species richness and composition of fruiting plants changes through time Figure 3 Cumulative fruiting curves calculated starting from 1997 and 1998 fruiting peaks Over 70 of species fruit within a year Fruit Color and Size Compared to rain forests in Asia Africa and South America frugivorous birds and bats are restricted to a few species in Madagascar and fruit eating mammals are predominantly primate species Fleming 1987 With a lower diversity of seed dispersers than other geographic regions one could expect plants to produce fruits of relatively uniform size and color For example extant seed dispersing mammals bats and birds in Madagascar’s rain forest are smaller in body size than those in African Asian and South American forests Fleagle amp Reed 1995 Godfrey et al 1997 and smaller fruit size would be expected Indeed only 5 out of the 98 species studied in this work had fruits larger than 50g Color vision capabilities also vary greatly between continental areas In Asia and Africa primates have excellent color vision Dominy 2001 in South America most monkeys have color vision but are deficient in red wave lengths Jacobs 1993 and Madagascar’s lemurs have been shown to be largely devoid of the physiological equipment necessary for color vision Jacobs 1993 Indeed in South America Janson found evidence that monkeys are attracted by the color and size of the fruit Janson 1983 Following this line of thinking we might predict that fruits would be less colorful in Madagascar where the color vision deficient lemurs are the predominant seed dispersers Of 87 species with data on fruit morphology 31 had red-orange 24 had light yellow or white 21 had green and 11 had dark coloration This suggests that plants could have coevolutionary relationships with specific seed dispersers or alternatively that fruit colors could bear little adaptive significance in Madagascar At least understory plants with dark fruits such as Psychotria may rely on birds as seed dispersers This is indicated by the fact that the smaller size of dark fruits which have an average weight of 152 g compared to 42 – 67 g average weights of fruits in the other color groups While all of the fruit color groups have synchronous fruiting through time Fig 4 sifakas appear to prefer red-orange fruits over other colors Sifakas ate 14 45 of the red-orange 8 33 of the light and 8 38 of the green species No dark species were eaten Figure 4 Fruit color and phenology of fruit availability Note how all fruit colors fluctuate synchronously in the number of species carrying fruit This putative fruit color preference in sifakas fitting the fact that Ranomafana forest has fruits of different colors is corroborated by recent color discrimination tests showing that some lemurs can distinguish colors quite well Gosset amp Roeder 2000 Jacobs amp Deegan 1993 One possible explanation is that even with only two classes of cone pigments dichromatic their behavioral performance can result from an ability to use signals from rods and cones jointly Jacobs amp Deegan 1993 Sifakas showed also a slight preference towards larger fruits Of the fruiting species that had fruits heavier than 1g 16 39 species were eaten by sifakas while 14 30 of lt1g species were eaten Species with larger fruits showed earlier seasonal fruiting peaks Fig 5 Correlation between large and small fruited species is 070 Spearman rank correlation P 0002 when fruiting of large fruits is shifted one month later Correlation without temporal correction is 044 P 0027 suggesting that there is at least a one month delay in the fruiting peak of species with small fruits Fig 5 Figure 5 Fruit size and phenology of fruit availability Note how species with large fruits peak before species with small fruits Lemur Response to Scarce Food Periods The seasonal tracking of fruit feeding by sifakas Figs 1B 2 which reflects both the diversity and amount of available fruit indicates that this lemur species is not dependent on fruits for individual survival during the lean season In the case of sifakas the drop in fruit feeding corresponds to an increase in leaf eating In addition to dietary shifts all lemur species appear to be able to deal with the season of scarce resources by conserving energy Ganzhorn 1993 Morland 1993 Nash 1998 Schmid 1998 Schmid amp Ganzhorn 1996 Wright amp Martin 1995 Extreme responses to winter season are seen in small-bodied lemurs Cheirogaleus spp go into hibernation for 4-6 months every year Muller 1998 Wright amp Martin 1995 and Microcebus also enter torpor for several days at a time Atsalis 1999 Fietz 1998 Schmid 1998 Lepilemur ruficaudatus has the lowest basal metabolic rate recorded for any folivorous mammal Schmid amp Ganzhorn 1996 Additional lemur traits that can promote energy conservation are thick insulating fur increased resting behavior maintenance of small group size birth of low-weight infants and relatively small brain size Wright 1999 The seasonal shift in sifaka diet does not however indicate that these lemurs are completely able to replace fruits with leaves in terms of energy Sifakas lose up to 20 of their weight during winter season Pochron amp Wright 2002 PCS unpubl data suggesting that fruits play an important role even in this relatively folivorous lemur At least 30 of sifaka feeding time is on fruits annually while species of the most frugivorous lemur genera Eulemur rubriventer and Varecia variegata spend 70 and 90 of their annual feeding time on fruits respectively Balko 1998 Overdorff 1991 Varecia a highly arboreal lemur weighing three to four kg can be regarded as the only living lemur that is an obligate frugivore It is also relatively specialized because fruits from five species make up to two thirds of its annual diet Balko 1998 This specialized frugivory is reflected in Varecia having large territories and extreme seasonal shifts in territory use Balko 1998 However even Varecia has been reported to survive on leaves after total loss of fruit productivity due to cyclone damage on trees albeit with a substantial weight loss and lack of reproduction Ratsimbazafy 2002 Ratsimbazafy et al in press Fruits As Keystone Resources for Reproductive Output Individual lemur species have strict breeding synchrony with a mating season typically lasting less than two weeks Rasmussen 1985 Sauther 1991 This breeding synchrony is triggered by changes in photoperiodicity which makes lemurs uniquely coupled with seasonal changes among primates Pereira 1993 van Horn 1975 Furthermore in contrast to most primate communities in the Neotropics Africa or in Asia Chapman et al 1999 Gautier-Hion et al 1985 Struhsaker 1997 Terborgh 1983 lemurs do not have synchronous birth peaks across species In Ranomafana sympatric lemurs show that while individuals within a species have synchronized births different species gave birth at different times of year Wright 1999 Fig 6 However while mating and birth seasons are not synchronized across lemur species weaning appears to happen in all species during March-April Fig 6 Figure 6 Reproductive schedules for 9 of 12 species of the sympatric lemur community at Ranomafana National Park Note birth asynchrony and weaning synchrony in these rain forest species Weaning occurs around the end of March For information about individual species see Table 1 One effect of the weaning synchrony is that all lemur species lactate during the period of increasing fruit availability Fig 7 The smallest lemurs are able to fit their whole breeding cycle into the peak fruiting season while sifakas the largest of the living lemurs at up to 7kg lactate for two first months without fruits Figs 2 6 7 It is noteworthy that while lactation is the most energy demanding stage of reproduction Lee 1997 Tilden amp Oftedal 1997 sifaka newborns are small relative to their mothers A newborn sifaka weights around 100g which is less than 2 of the mother’s weight and thus the initial cost of lactation is far less than during the peak fruiting season when the infant is over 15 of mother’s weight Wright 1999 The peak lactation synchrony among sympatric lemurs Fig 7 suggests that even sifakas while relatively opportunistic fruit eaters Figs 1 2 may rely on fruits as key resources for reproductive success Figure 7 Percentage of plant species fruiting lemur species gestating and lemur species lactating Note how lactation and fruiting peaks coincide while largest number of lemurs gestate during austral winter months when the fruit availability is low It is also interesting that bigger fruits peak in abundance prior to smaller fruits in the forest Fig 5 The delayed peak richness of small fruits coincides with the beginning of weaning period in lemurs Weaning marks a nutritional transition when the infant becomes a juvenile foraging independently and is more vulnerable to the risks of malnutrition infection and predation Janson amp van Schaik 1993 For plants this synchrony of weaning produces a population peak of small juvenile lemurs foraging in the forest for fruits effectively resulting in a peak of seed dispersers Even bamboo-eating lemurs genus Hapalemur follow the weaning synchrony and include small amounts of fruit to their diet during the summer fruiting season Tan 1999 Grassi 2001 The peak fruit production in the Malagasy rain forest is three months shorter on average compared to peak fruit production in the Amazon forest Stevenson this volume Terborgh 1983 Wright 1997b and the African forest Chapman this volume Gautier -Hion et al 1985 Struhsaker 1997 This suggests that the environment in Madagascar has more well-defined constraints than other continental areas where primates have evolved In this respect it is informative that lemurs do not appear to rely on fruits to carry them over the period of fruit scarcity as would be expected from classical descriptions of keystone resources Terborgh 1983 Soil fertility overall in Madagascar is low often lower than in other primate habitats on other continents Ganzhorn et al 1999 Smith and Ganzhorn 1996 compared lemurs and Australian marsupials suggesting that both radiations were strongly influenced by these restricting environmental factors that contrast with many tropical habitats in South America Africa and mainland Asia Thus even if lemur taxa had evolved to use classical keystone resources to survive winter seasons the harsher environmental constraints of Madagascar may have weeded them out In conclusion the data presented in this paper suggest that lemurs as a guild rely on fruits as a keystone resource during the warm wet months in order for lactation and weaning to succeed It is important to note that many of the fleshy-fruited plant species used by lemurs are also hardwood species favored by selective loggers While loss of key fruit trees may not drive lemurs into extinction immediately it may adversely affect reproductive success years after logging Pochron et al 2004 This kind of extinction debt might remain unnoticed if keystone resource is considered in forest management and reserve planning only in the narrow sense Thus keystone resources are probably not just the ones that help ecologically important animal guilds survive a bad season ACKNOWLEDGEMENTS In Madagascar we would like to thank the National Association for the Management of protected Areas ANGAP Department of Water and Forests and the Minister of the Environment We also thank Benjamin Andriamihaja and the MICET and ICTE staff who expedite the research process and have made our long-term work in Madagascar possible We acknowledge the David and Lucile Packard Foundation Douroucouli Foundation the Wenner-Gren Foundation the John D and Catherine T MacArthur Foundation National Geographic Society National Science Foundation USA Earthwatch Institute and SUNY-Stony Brook We would also like to acknowledge Paul Rasabo the late George Rakotonirina Remi Rakotovao Raymond Ratsimbazafy Pierre Talata and Albert Telo for their expert assistance following animals and collecting phenological data Our thanks to Ted Stiles who inspired this project and assisted with setting up the methods References Abraham J-P Rakotonirina B Randrianasolo M Ganzhorn JU Jeannoda V Leigh EG Jr 1996 Tree diversity on small plots in Madagascar: a preliminary review Rev Ecol 51:93-117 Atsalis S 1999 Diet of the brown mouse lemur Microcebus rufuss in Ranomafana National Park Madagascar Int J Primatol 20: 193-229 Balko EA 1998 A Behaviorally Plastic Response to Forest Composition and Logging Disturbance By Varecia variegata variegata In Ranomafana National Park Madagascar State University of New York-Syracuse: Syracuse Bawa KS Ashton PS and Nor SM 1990 Reproductive ecology of tropical forest plants: management issues In KS Bawa and M Hadley ede Reproductive ecology of tropical forest plants Parthenon Carnforth UK Chapman CA Gautier-Hion A Oates JF and Onderdonk D 1999 African primate communities: Determinants of structure and threats to survival In JG Fleagle CH Janson and K Reed eds: Primate Communities Cambridge: Cambridge University Press pp1-37 Dew JL and Wright PC 1998 Frugivory and seed dispersal by four species of primates in Madagascar’s eastern rainforest Biotropica 30:1-13 Dominy NJ and Lucas PW 2001 Ecological importance of trichromatic vision to primates Nature 410: 363-366 Fietz J 1998 Body mass in wild Microcebus murinus over the dry season Folia Primatol 69: 183-190 Fleagle JG and Reed K 1996 Comparing primate communities: a multivariate approach J Hum Evol 30:489-510 Fleming TH Breitwisch R Whitesides GH 1987 Patterns of tropical frugivore diversity Ann Rev Ecol Syst 18: 91-109 Foster RB 1982 Famine on Barro Colorado Island In EG Leigh Jr AS Rand and DM Windsor eds: The Ecology of a Tropical Forest: Seasonal Rhythms and Long-term Changes Washington DC: Smithsonian Institution Press pp 201-212 Ganzhorn JU 1993 Flexibility and constraints of Lepilemur ecology In PM Kappeler and JU Ganzhorn eds: Lemur Social Systems and Their Ecological Basis New York: Plenum pp 153-165 Ganzhorn JU and Kappeler PM 1996 Lemurs in the Kirindy forest In JU Ganzhorn and JP Sorg eds: Ecology and Economy of a Tropical Dry Forest in Madagascar Gottingen:Erich Goltze GmbH amp Co pp 57-80 Ganzhorn JU Wright PC and Ratsimbazafy HJ 1999 Primate Communities: Madagascar In JG Fleagle CH Janson and K Reed eds: Primate Communities Cambridge: Cambridge University Press pp 75-89 Gautier-Hion A Duplantier JM Quiris F Feer R Sourd C Decoux JP Emmons L Erard C Heckestweiler P Moungazi A Roussilhon C and Thiollay JM 1985 Fruit characters as a basis of fruit choice and seed dispersal in a tropical forest vertebrate community Oecologia 65: 324-337 Gentry AH 1993 Diversity and Floristic composition of lowland tropical forest in Africa and South America in Goldblatt P Editor: Biological Relationships between Africa and South America Yale University: New Haven p 500-547 Gentry AH and Terborgh J 1990 Composition and dynamics of the Cocha Cashu “mature” floodplain forest In AH Gentry ed: Four Neotropical Rainforests New Haven: Yale University Press pp 542-564 Gilbert LE 1980 Food web organization and conservation of of neotropical biodiversity In ME Soulé and BA Wilcox eds: Conservation Biology pp 11-34 Sinauer Sunderland MA Glander KE Wright PC Merenlender A and Daniels PS 1992 Morphometrics and testicle size of rainforest lemur species from southeastern Madagascar J Hum Evol 22:1-17 Godfrey LR Jungers WS Reed KE Simons ELS and Chatrath PS 1997 Subfossil lemurs: Inferences about past and present Primate Communities in Madagascar In SM Goodman and BD Patterson eds: Natural Change and Human Impact in Madagascar Washington: Smithsonian Press pp 218-256 Goodman SM and Ganzhorn JU 1997 Rarity of figs Ficus on Madagascar and its relationship to a depauperate frugivore community Revue d’Ecologie-La terre et la Vie 52 4: 321-329 Gosset D and Roeder JJ 2000 Colour and shape discrimination in black lemurs Eulemur macaco Folia primatol 71: 173-176 Grassi C 2001 The behavioral ecology of Hapalemur griseus griseus: The influences of microhabitat and population density on this small-bodied prosimian folivore PhD University of Texas at Austin Hemingway CA 1995 Feeding and reproductive strategies of the Milne-Edward’s Sifaka Propithecus diadema edwardsi Unpubl PhD dissertation Duke University Hemingway CA 1996 Morphology and phenology of seeds and whole fruit eaten by Milne-Edward’s sifaka Propithecus diadema edwardsi in Ranomafana National Park Madagascar Int J Primatol 17: 637-660 Hemingway CA 1998 Selectivity and variability in the diet of Milne-Edwards’ sifakas Propithecus diadema edwardsi: Implications for folivory and seed-eating Int J of Primatol 19:355-377 Jacobs GH Deegan JF 1993 Pigments underlying color vision in ringtailed lemurs Lemur catta and brown lemurs Eulemur fulvus Am J Primatol 30: 243-256 Janson CH 1983 Adaptation of fruit morphology to dispersal agents in a neotropical forest Science 219: 187-189 Janson CH and van Schaik CP 1993 Ecological risk aversion in juvenile primates: slow and steady wins the race In ME Pereira and LA Fairbanks eds: Juvenile Primates Life History Development and Behavior Oxford: Oxford University Press pp 57-74 Krause DW Hartman JH and Wells NA 1997 Late Cretaceous vertebrates from Madagascar : Implications for biotic change in deep time In SM Goodman and BD Patterson eds: Natural Change and Human Impact in Madagascar Washington: Smithsonian Press pp 3-43 Kress WJ Schatz GE Andrianifahanana A and Morland HS 1992 Pollination of Ravenela madagascariensis by lemurs in Madagascar: evidence of an archaic coevolutionary system American Journal of Botany 81: 542-551 Langrand O 1990 The Birds of Madagascar Yale University Press New Haven Lee PC 1997 The meanings of weaning: growth lactation and life history Evol Anthropol 5:87-96 Lovejoy T and Bierregaard RO Jr 1990 Central Amazonia forests and the minimum critical size of ecosystems project In AH Gentry ed: Four Neotropical Rainforests New Haven: Yale University Press pp 60-74 Lowry PP II Schatz GE and Phillipson PB 1997 The classification of natural and anthropogenic vegetation in Madagascar In SM Goodman and BD Patterson eds: Natural Change and Human Impact in Madagascar Washington: Smithsonian Press pp 93-123 Mittermeier RA Tattersall I Konstant WR Meyers DM and Mast RB 1994 Lemurs of Madagascar Washington DC: Conservation International Morland HS 1991 Preliminary report on the social organization of ruffed lemurs Varecia variegata variegata in a Northeast Madagascar rainforest Folia Primatol 56:157-161 Morland HS 1993a Reproductive activity of ruffed lemurs Varecia variegata variegata in a Madagascar Rain Forest Am J Phys Anthropol 97:71-82 Morland HS 1993b Seasonal behavioral variation and its relationships to thermoregulation in ruffed lemurs In PM Kappeler and JU Ganzhorn eds: Lemur Social Systems and Their Ecological Basis New York: Plenum pp 193-205 Muller AE 1998 A preliminary report on the social organization of Cheirogaleus medius Cheirogalidae Primates in north-west Madagascar Folia Primatol 69: 160-166 Nash LT 1998 Vertical clingers and sleepers: Seasonal influences on the activities and substrate use of Lepilemur leucopus at Beza Mahafaly Special Reserve Madagascar Folia Primatol 69: 204-217 Nilsson LA Rabakoandrianina E Pettersson B and Grunmeier R 1993 Lemur pollination in the Malagasy rainforest liana Strongylodon craveniae Leguminosae Evolutionary Trends in Plants 7: 49-56 Nunes A 1998 Diet and feeding ecology of Ateles belzebuth belzebuth at Maraca Ecological Station Roraima Brazil Folia Primatol 69: 61-76 Oates JF Whitesides GH Davies AG Waterman PG Green SM Dasilva GL and Mole S 1990 Determinants of variation in tropical forest primate biomass: new evidence from West Africa Ecology 71: 328-343 Overdorff DJ 1991 Ecological correlates to social structure in two prosimian primates: Eulemur fulvus rufus and Eulemur rubriventer in Madagascar Overdorff DJ 1992 Differential patterns in flower feeding by Eulemur fulvus rufus and Eulemur rubriventer in Madagascar Am J Phys Anthropol 28:191-203 Overdorff DJ 1993 Similarities differences and seasonal patterns in the diets of Eulemur rubriventer and Eulemur fulvus rufus in the Ranomafana National Park Madagascar Int J Primatol 14:721-753 Overdorff DJ and Strait SG 1998 Seed handling by three prosimian primates in southeastern Madagascar: implications for seed dispersal Am J Primatol Pereira ME 1993 Seasonal adjustment of growth rate and adult body weight in ringtailed lemurs In PM Kappeler and JU Ganzhorn eds Lemur Social Systems and Their Ecological Basis New York Plenum pp 205-221 Peres C 1993 Sturcture and spatial organization of an Amazonian terra firme forest primate community J of Tropical Ecology 9: 259-276 Peterson RL et al 1995 Chiroptères Faune de Madagascar 84:1-204 Pochron ST and Wright PC 2002 Dynamics of testes size compensates for variation in male body-size Evolutionary Ecology Research 4: 577-585 Pochron ST and Wright PC 2004 Flexibility in adult group compositions of a prosimian primate Behavioral Ecology and Sociobiology in press Pochron ST Tucker WT and Wright PC 2004 Demography life history and social structure in Propithecus diadema edwardsi from 1986 to 2000 of Ranomafana National Park Madagascar American Journal of Physical Anthropology in press Powzyk JA 1997 The Socio-Ecology of Two Sympatric Indriids: Propithecus diadema diadema and Indri indri A Comparison of Feeding Strategies and Their Possible Repercussions on Species-Specific Behaviors unpubl PhD Dissertation Duke University Durham NC Razafindratsita VM 1995 The feeding ecology of the velvet asity Philapetta DEA University of Antanarivo Madagascar Rasmussen T 1985 A comparative study of breeding seasonality and litter sizein eleven taxa of captive lemurs Lemur and Varecia Int J Primatol 6: 501-511 Ratsimbazafy HJ 2002 On the Brink of Extinction and the Process of Recovery: Varecia variegata variegata in Manombo Forest unpubl PhD dissertation State University of New York Stony Brook Ratsimbazafy HJ Jungers W Louis E and Wright PC in press Reproductive success in Varecia variegata in Manombo Reserve: recovery from a cyclone J of Hum Ev Reitsma JM 1988 Vegetation forestiere du Gabon Tropenbos Technical Series 1:1-142 Sauther ML 1991 Reproductive behavior of free-ranging Lemur catta at Beza Mahafaly Special Reserve Madagascar Am J Phys Anthropol 84:463-477 Schatz GE Lowry PP II and Phillipson PB 2002 The Trees of Madagascar Missouri Botanical Garden Press St Louis Schmid J 1998 Daily torpor in mouse lemurs Microcebus spp: Metabolic rate and body temperature Folia Primatol 69: 394-404 Schmid J and Ganzhorn JU 1996 Resting metabolic rates of Lepilemur ruficaudatus Am J Primatol 38:169-174 Simons EL 1997 Lemurs old and new In SM Goodman and BD Patterson eds: Natural Change and Human Impact in Madagascar Washington: Smithsonian Press pp 142-166 Smith AP and Ganzhorn JU 1996 Convergence and divergence in community structure and dietary adaptations in Australian possums and gliders and Malagasy lemurs Australian J of Ecol 21: 31-46 Struhsaker TT 1997 Ecology of an African Rain Forest: Logging in Kibale and the Conflict between Conservation and Exploitation Gainesville: University Press of South Florida Sussman RW and Rakotozafy A 1994 Plant diversity and structural analysis of a tropical dry forest in southwestern Madagascar Biotropica 26:241-254 Tan CL 1999 Group composition home range size and diet of three sympatric bamboo lemur species genusHapalemur in Ranomafana National Park Madagascar Int J of Primatol20:547-566 Tattersall I 1982 The Primates of Madagascar New York: Columbia University Press Terborgh J 1983 Five New World Primates: A Study in Comparative Ecology Princeton:Princeton University Press Terborgh J 1986 Keystone plant resouces in the tropical forest In Soule M ed Conservation Biology: The Science of Scarcity and Diversity Sunderland Sinauer Ass Incpp 330-344 Tilden CD and Oftedal OT 1997 Milk composition reflects pattern of maternal care in prosimian primates Am J Prinatol 41: 195-211 Van Horn RN 1975 Primate breeding season: photoperiodic regulation in captive Lemur catta Folia primatologica 24: 203-212 Van Schaik CP 1986 Phenological changes in a Sumatran rain forest J Trop Ecol 2:327-247 Wright PC 1989 The nocturnal primate niche in the New World J Hum Evol 18:635-646 Wright PC 1992 Primate ecology rainforest conservation and economic development: Building a national park in Madagascar Evol Anthropol 1:25-33 Wright PC 1995 Demography and life history of free-ranging Propithecus diadema edwardsi in Ranomafana National Park Madagascar Int J Primatol 16:835-854 Wright PC 1997a The future of biodiversity in Madagascar: a view from Ranomafana National Park In SM Goodman and BD Patterson eds: Natural Change and Human Impact in Madagascar Washington: Smithsonian Press pp 381-405 Wright PC 1997b Behavioral and ecological comparisons of Neotropical and Malagasy primates In WG Kinzey ed New World Primates: Ecology Evolution and Behavior New York: Aldine de Gruyter pp 127-143 Wright PC 1998 Impact of predation risk on the behaviour of Propithecus diadema edwardsi in the rain forest of Madagascar Behaviour 135:1-30 Wright PC 1999 Lemur traits and Madagascar ecology: coping with an island environment Yearbook of Physical Anthropology 42: 31-72 Wright PC and Martin LB 1995 Predation pollination and torpor in two nocturnal primates: Cheirogaleus major and Microcebus rufus in the rainforest of Madagascar In L Alterman G Doyle and MK Izard eds: Creatures of the Dark New York: Plenum Press pp 45-60 Wright P C and Andriamihaja BA 2002 Making a rain forest national park work in Madagascar: Ranomafana National Park and its long-term research commitment In J Terborgh C van Schaik M Rao and L Davenport eds: Making Parks Work: Strategies for Preserving Tropical Nature California: Island Press pp 112-136 Dept of Anthropology SUNY Stony Brook Stony Brook New York USA Institute for the Conservation of Tropical Environments SUNY Stony Brook Stony Brook New York USA Centre VALBIO Ranomafana National Park Madagascar Institute for the Conservation of Tropical Environments Antananarivo 101 Madagascar and Centre VALBIO Ranomafana National Park Madagascar Department of Anthropology SUNY Stony Brook Stony Brook New York USA Institute of Biotechnology University of Helsinki Helsinki Finland |
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